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It was long thought that maize was domesticated solely from parviglumis --- a subspecies of teosinte, a wild grass --- in Mexico's lowlands about 9,000 years ago. However, a 2024 analysis has revealed that between 15 and 25percent of maize's genes can be traced to a second ______ highland subspecies of teosinte that was hybridized with maize around 4,000 years after the initial domestication.