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Verbal·Standard English Conventions·Boundaries

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The relationship between grassland arthropod populations and agricultural fertilizers containing macronutrients—such as nitrogen, phosphorus, and potassium—was the focus of a 2000 study in the __________ by the researcher Mark E. Ritchie, the study examined a variety of arthropod specimens, which had been collected with sweep nets.

Which choice completes the text so that it conforms to the conventions of Standard English?

A

United States led

B

United States, led

C

United States. Led

D

United States and led