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Some scientists have suggested that as ocean temperatures rise, many fish and corals found in the shallow zone (less than 30 meters below the surface) could take refuge in the cooler, darker mesophotic zone (30 to 150 meters below the surface). However, it isn't clear that all such species will be able to tolerate mesophotic conditions. In 2018, Hudson Pinheiro and colleagues studied stony corals in the two zones in Caribbean and Indo-Pacific waters. Based on the depths at which those corals are now found, the species that seems least suited to a full migration to the mesophotic zone is _____