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Verbal·Standard English Conventions·Boundaries

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Biochemist Jennifer Doudna, along with her French collaborator Emmanuelle        the CRISPR-Cas9 microbial adaptive immune system into a programmable genome-editing technology that revolutionized biomedical science.

Which choice completes the text so that it conforms to the conventions of Standard English?

A

Charpentier, adapted

B

Charpentier adapted

C

Charpentier; adapted

D

Charpentier—adapted